Master of Science (MS), Ohio University, 2023, Geological Sciences
Studying extant tracemakers, environmental conditions, trace construction, and trace morphology is vital for ichnofossil interpretation. Many scorpions are known to burrow, but despite having existed since the Silurian, they have a poor trace fossil record likely due to a lack of understanding of modern scorpion traces. In this study, burrows of three extant scorpions, Heterometrus spinifer a tropical species from Asia, Pandinus viatoris a tropical species from Africa, and Paravaejovis spinigerus an arid species from North America, were studied to identify key burrow characteristics, determine if and how evolutionary distance changed burrow form, and determine how burrow construction and form was affected by sediment properties. Neoichnological experiments were used to determine the burrowing techniques, behaviors, and trace morphologies of each species under natural conditions and under conditions of altered soil moisture and composition.
Scorpions were placed in 246-, 212-, 114-, or 38-liter terrariums based on scorpion size for three experiments, each lasting 14-30 days. The experiments consisted of 1) substrate and moisture conditions based on natural conditions, 2) raising or lowering moisture content from natural conditions, and 3) increasing or decreasing sediment grain size from natural conditions. At the end of each experiment, the scorpions were removed and burrows were cast in plaster for qualitative and quantitative description and analysis.
Burrows produced in this study bore a moderate-to-high similarity to each other despite differences in excavation methods and architecture. Heterometrus spinifer and Pandinus viatoris used similar excavation methods where sediment was gathered in the first two pairs of legs and/or chelicerae, dragged away, and dropped. Heterometrus spinifer produced open, straight burrows comprised of a single entrance and subvertical tunnel. Pandinus viatoris produced large diameter, sinuous, branching burrow networks with 1-3 ent (open full item for complete abstract)
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Committee: Greg Springer (Advisor); Katherine Fornash (Committee Member); Daniel Hembree (Committee Member)
Subjects: Biology; Ecology; Environmental Studies; Experiments; Geology; Paleoecology; Paleontology; Zoology